Light Apple Muffins

Ingredients

• 1 cup all purpose flour
• 1 cup whole wheat flour
• 1 cup wheat germ
• 1/4 cup firmly packed brown sugar
• 1 teaspoon baking powder
• 1 teaspoon baking soda
• 1/4 teaspoon salt
• 1 teaspoon ground cinnamon
• 1/4 teaspoon ground nutmeg
• 2 eggs beaten
• 1/4 cup skim milk
• 1/4 cup vegetable oil
• 2 cups apple peeled, shredded
• 1 tablespoon grated lemon rind
• cooking spray